Pro-Vice Chancellor for Medicine and Health Sciences , University of East Anglia
Former Head, Division of Biological Sciences , University of Edinburgh
Emeritus Professor of Pathology , University of Cambridge
Regius Professor of Life Sciences , University of Dundee
Consulting Professor of Biochemistry , University of Dundee
Professor & Wellcome Principal Research Fellow , University of Edinburgh
Professor of Haematological Regeneration , University of Edinburgh
Former Professor of Biochemistry , Dynamic Earth Enterprises Ltd
Emeritus Professor of Bacterial Genetics , University of Edinburgh
Professor of Cell Biology , University of Glasgow
Director, Australian Regenerative Medicine Institute , Monash University
Former Head, Imperial Cancer Research Fund Tumour Virus Group , University of Cambridge