Professor of Molecular Biology , University of Dundee
Director, MRC Human Genetics Unit , Western General Hospital
Professor in Cellular Signalling , University of Dundee
Former Robertson Professor of Bioprocess Technology , University of Strathclyde
Deputy Vice-Chancellor (Research and Impact) , University of Exeter
Research Professor of Biology and Biological Engineering , California Institute of Technology
Researcher , Universidad de la Laguna
Former Non-Executive Director and Chairman , City, University of London
Professor of Plant Biochemistry , University of Edinburgh
Professor, Department of Zoology , University of Texas at Austin
Professor of Biochemistry , University of Aberdeen
Formerly Group Leader , Western General Hospital