Emeritus Professor of Anatomy , University of Edinburgh
Emeritus Professor of Biochemistry , University of Aberdeen
Programme Leader , Roslin Institute
Programme Leader, MRC Human Genetics Unit , University of Edinburgh
Member, International Advisory Board, Chinese Academy of Sciences , Chinese Academy of Sciences
Professor of Mathematical & Computational Cell Biology , University of Glasgow
Emeritus Group Leader , Francis Crick Institute
Professor of Structural Biology , University of Dundee
Deputy Director, Wellcome Centre for Anti-Infectives , University of Dundee
Professor of Molecular Microbiology , University of Strathclyde
Former Pro-Vice-Chancellor of Science, Agriculture and Engineering , Newcastle University
Former Fellow, Hertford College , Durham University