Professor of Molecular Biology , University of Dundee
(Deceased) Emeritus Professor of Biological Timing , University of Edinburgh
Professor of Molecular Genetics , University of Edinburgh
Former Sibthorpian Professor of Plant Science , University of Oxford
Professor of Genome Integrity , University of Dundee
Professor of Chemistry and Bioengineering , Massachusetts Institute of Technology
Past President , Royal Society
Former Professor of Molecular Cell Biology , University College Dublin
Emeritus Professor of Molecular Parasitology , University of Edinburgh
UK Adviser in Biotechnology , Remedios Ltd