Professor of Biophysics, ETH Zürich.

Emeritus Professor, Newcastle University.

Professor of Reproductive Science, University of Edinburgh.

Former Chief Executive Officer, Horticulture Research International, Wellesbourne.

Hansen Professor of Structural Biology, The Scripps Research Institute.

Professor of Biochemistry, University of St Andrews.

Professor of Biochemistry, University of York.

Former Professor of Developmental Biology, University of Dundee.

Professor of Crop Genomics, James Hutton Institute.

Professor of Immunobiology, University of Dundee.

Professor of Developmental Physiology, University of Dundee.

Chief Scientist and Group Leader, Francis Crick Institute, Cancer Research UK.