(Deceased) Professor Emeritus , University of Aberdeen
Honorary Fellow and Emeritus Professor of Zoology , University of Edinburgh
Regius Professor of Zoology , University of Glasgow
Emeritus Professor of Forestry , University of Aberdeen
Professor of Behavioural Ecology , University of Glasgow
Former Reader in Agricultural Biochemistry , University of Edinburgh
Former Director, Dunstaffnage Marine Laboratory
Former Director , Macaulay Land Use Research Institute
Former Chief Scientific Adviser to the UK Government , University of Oxford
Visiting Professor of Freshwater Ecology , Fish Conservation Centre
Distinguished Research Professor of Mathematical Biology , University of California, Santa Cruz
Professor Emeritus of Natural History , University of Edinburgh