Former Regius Professor of Botany, University of Edinburgh.

Honorary Curator of Entomology, Harvard University.

Head of Conservation Science, RSPB Scotland.

Former Director, Royal Zoological Society of Scotland.

Former Deputy Director, AFRC Animal Breeding Research Organisation, Agricultural and Food Research Council (AFRC).

Emeritus Regius Professor of Botany, University of Glasgow.

Professor Emeritus of Agriculture and Rural Economy, University of Edinburgh.

Emeritus Professor of Microbiology, University of Glasgow.

Former Head of Mycology and Plant Pathology, Royal Botanic Garden Edinburgh.

Emeritus Professor, University of Nottingham.

Research Ecologist, Centre for Ecology and Hydrology.

Honorary Professor, University of Stirling.