Former Director, Beatson Institute for Cancer Research.

Emeritus Professor of Experimental Haematology, University of Dundee.

Former Dean, Faculty of Veterinary Medicine, University of Glasgow.

Emeritus Professor of Developmental Disabilities in Childhood, University of Edinburgh.

Professor of Molecular Pharmacology, University of Dundee.

Chair of Infectious Disease Epidemiology, University of Edinburgh.

MRC External Scientific Staff, University of Edinburgh.

Emeritus Wardlaw Professor of Evolutionary and Developmental Psychology, University of St Andrews.

Director of Liverpool School of Art & Design, Liverpool John Moores University.

Professor of Medical and Veterinary Molecular Epidemiology, Director, Global Health Academy and Vice-Principal Global Access, University of Edinburgh.

Professor of Psychology, University of Stirling.