Principal and Vice-Chancellor, Heriot-Watt University
Former Vice-President, Research Strategy Worldwide, Merck Research Laboratories
Wykeham Professor of Logic, University of Oxford
Professor of Neurology, University of Glasgow
Honorary Professor, University of Edinburgh
Gregory Chair of Applied Mathematics, University of St Andrews
Charlotte Street Partners
Former Chairman, Scottish Biomedical Foundation
Chair, Department of Integrative Structural and Computational Biology and Hansen Professor of Structural Biology, The Scripps Research Institute
Professor of Human Genetics, University of Edinburgh
Honorary Research Fellow, James Hutton Institute